Chain A ( Y293 T302 V305 H306 Y308 V309 G310 K311 E312 P313 S314 G315 L316 R317 F318 N319 K320 L321 F322 L323 N324 E325 D326 D327 K328 P329 H330 N331 P332 M333 V334 N335 A336 G337 R387 Y394 K398 L459 H461 S462 C463 G464 M465 Y466 D467 F468 S469 G470 Q471 F472 A473 F474 H475 K481 S482 G483 M508 N510 ),  Chain C ( K289 P290 Y293 T302 V305 H306 Y308 V309 G310 K311 E312 P313 S314 F318 N319 K320 L321 F322 L323 N324 E325 D326 D327 K328 P329 H330 N331 P332 M333 V334 N335 A336 G337 R387 Y394 K398 C400 S458 L459 M460 H461 S462 C463 G464 M465 Y466 D467 F468 S469 G470 Q471 F472 A473 F474 H475 A480 K481 S482 G483 G487 I488 M508 N510 ),  Chain B ( K320 L321 F322 L323 N324 E325 D326 D327 R387 Y394 K398 ),  Chain D ( R317 F318 K320 L321 F322 L323 E325 D327 R387 Y394 K398 )